As a puppy, Bella finds her way into the arms of Lucas, a young man who gives her a good home. When Bella becomes separated from Lucas, she soon finds herself on an epic, 400-mile journey to reunite with her beloved owner. Along the way, the lost but spirited dog touches the lives of an orphaned mountain lion, a down-on-his-luck veteran and some friendly strangers who happen to cross her path.

Dean is an illustrator whose unwillingness to deal with the recent death of his mother means escaping his hometown of New York for an interview with an ad agency in Los Angeles. His retired engineer dad Robert takes a more regimented approach to grief, including putting the family home up for sale. Both father and son set out on their own paths to find a new normal as unexpected circumstances and potential new love interests threaten to thwart all...